Baha El Din, S.M. (2007) - A new lizard of the Acanthodactylus scutellatus group (Squamata: Lacertidae) from Egypt. - Zoology in the Middle East, Heidelberg, 40 (1): 21-32.   × Es warden zwei morphologisch und ökologisch unterschiedliche Populationen von Eidechsen aus der Acanthodactylus scutellatus-Artengruppe dokumentiert, die bisher als konspezifisch unter dem Namen A. longipes behandelt wurden. Es wird gezeigt, dass beide Populationen in einem großen Arealbereich sympatrisch vorkommen. Die Zugehörigkeit von Tieren aus der Westliche Wüste Ägyptens zu A. longipes wird bestätigt, während Populationen aus östlichen Landesteilen, die bisher ebenfalls als zu A. longipes gehörend angesehen wurden, als neue Art beschrieben wird. Tamar, K. & Carranza, S. & Sindaco, R. & Moravec, J. & Trape, J.-F. & Meiri, S. (2016) - Out of Africa: Phylogeny and biogeography of the widespread genus Acanthodactylus (Reptilia: Lacertidae). - Molecular Phylogenetics and Evolution, 103: 6-18. × Acanthodactylus lizards are among the most diverse and widespread diurnal reptiles in the arid regions spanning from North Africa across to western India. Acanthodactylus constitutes the most species-rich genus in the family Lacertidae, with over 40 recognized species inhabiting a wide variety of dry habitats. The genus has seldom undergone taxonomic revisions, and although there are a number of described species and species-groups, their boundaries as well as their interspecific relationships are largely unresolved. We constructed a multilocus phylogeny, combining data from two mitochondrial (12S, cytb) and three nuclear (MC1R, ACM4, c-mos) markers for 302 individuals belonging to 36 known species, providing the first large-scale time-calibrated molecular phylogeny of the genus. We evaluated phylogenetic relationships between and within species-groups, and assessed Acanthodactylus biogeography across its known range. Acanthodactylus cladogenesis is estimated to have originated in Africa due to vicariance and dispersal events from the Oligocene onwards. Radiation started with the separation into three clades: the Western and scutellatus clades largely distributed in North Africa, and the Eastern clade occurring mostly from Arabia to south-west Asia. Most Acanthodactylus species diverged during the Miocene, possibly as a result of regional geological instability and climatic changes. We support most of the current taxonomic classifications and phylogenetic relationships, and provide genetic validity for most species. We reveal a new distinct blanfordii species-group, suggest new phylogenetic positions (A. hardyi, A. masirae), and synonymize several species and subspecies (A. lineomaculatus, A. boskianus khattensis and A. b. nigeriensis) with their phylogenetically closely-related species. We recommend a thorough systematic revision of taxa exhibiting high levels of intraspecific variability as well as clear evidence of phylogenetic complexity such as A. guineensis, A. grandis, A. dumerilii, and A. senegalensis and the pardalis and erythrurus species-groups.
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